Here's some advice on that subject:
http://www.advrider.com/forums/showthread.php?t=287971
The main points:
- Bigger map screen resolution shows more map details, more interesting trails, fire roads etc. for the gravel-adventurers
- Better track handling capabilities
- Better user configuration possibilities - you can tailor-make your own user interface/display more than the newer, simplified units
- the x76,x78 series are primarily made for marine usage, where trees and other terrain don't interfere with sat reception
- the x76, x78 series have older technology ( less sensitive sat-reception, more expensive proprietary memory modules )
For offroad/gravel you may want better oversight over potential blind ends and zoom out to greater scales than the ordinary tarmac/main road travellers - thus making the x76/x78 series the better choice. The Zumo 550 has a different customer base targeted - smaller screen and less offroad capable with less user config possibilities. For example autozoom cannot be turned on/off like you can on the x76/x78 series.