I used to work on forecourts a lot, on the equipment side not the technical side of making fuel, and I'm pretty sure, although not certain, that I heard talk of different fuel compositions in the winter and summer.
I think it was to do with the amount of LPG blended back into the fuel (as LPG is by-product of petrol production, they want to get shot of it).
Also petrol is more dense in the winter, so 20 litres of fuel in the winter has more calories than 20 litres in the summer - but of course that would produce the oposite effect.
However in the underground storage tank it tends to vary by only a few degrees from mid winter to mid summer (the tank depth averages about 10 feet - that's 3m for those too young to remember VHS), but of course a bike tank is VERY exposed - so you fill the tank with toasty petrol, and then refrigerate it during your journey, hence it shrinks.